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Athyrium
Family: Athyriaceae
Athyrium image
Patrick Alexander
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Masahiro Kato in Flora of North America (vol. 2)
Plants generally terrestrial. Stems short-creeping or ascending, stolons absent. Leaves monomorphic, usually dying back in winter. Petiole ± 0.5 times length of blade or less, base swollen and dentate, persisting as trophopod over winter or not; vascular bundles 2, lateral, lunate in cross section. Blade lanceolate to elliptic or oblanceolate, 1--3-pinnate-pinnatifid, gradually reduced distally to confluent, pinnatifid apex, herbaceous. Pinnae not articulate to rachis, segment margins serrulate or crenate; proximal pinnae often reduced, sessile to short-petiolulate, ± equilateral; costae adaxially grooved, grooves continuous from rachis to costae to costules; indument absent or of linear to lanceolate scales or 1-celled glands abaxially. Veins free, simple or forked. Sori in 1 row between midrib and margin, round to elongate, straight or hooked at distal end, or horseshoe-shaped; indusia shaped like sori, persistent, attached laterally or with narrow sinus, or indusia absent. Spores brownish, rugose. x = 40. In species outside the flora stems are sometimes long-creeping to erect, with leaves radially or dorsiventrally arranged.

Vascular plants of NE US and adjacent Canada
Petiole stramineous, stout, with 2 bundles at the base, these uniting upwardly into a single trough-shaped bundle; blade pinnate to tripinnate, sparingly scaly, otherwise glabrous or sparsely hairy; veins free, simple or forked, reaching the margin; indusia borne on the anterior side of the vein, often recurved and crossing the vein and thus hooked or horseshoe-shaped, rarely back to back, subentire or toothed, often ciliate; large wood-ferns with thick, creeping to suberect rhizomes and thin, light green, deciduous lvs, the rhizome and often also the petioles with conspicuous, entire, fibrous scales; rhizome in our spp. beset with old swollen, starch-filled petiole-bases. 150+, cosmop.

Gleason, Henry A. & Cronquist, Arthur J. 1991. Manual of vascular plants of northeastern United States and adjacent Canada. lxxv + 910 pp.

©The New York Botanical Garden. All rights reserved. Used by permission.
Species within inventory project: North American Species Lists
Athyrium angustum
Image of Athyrium angustum
Athyrium asplenioides
Image of Athyrium asplenioides
Athyrium distentifolium
Image of Athyrium distentifolium
Athyrium filix-femina
Image of Athyrium filix-femina

The PCC, and this data portal, were made possible by funding from the National Science Foundation’s Advancing Digitization of Biological Collections (ADBC) program, grant numbers 1802504, 1802352, 1802134, 1802033, 1802270, 1802255, 1802239, 1802446, 1802305.
The pteridoportal taxonomic thesaurus is based on the Checklist of Ferns and Lycophytes of the World, generously provided by Michael Hassler.